[1] Jackwood M W. Review of infectious bronchitis virus
around the world [J]. Avian Dis, 2012, 56(4): 634-41.
[2] Cavanagh D. Coronavirus avian infectious
bronchitis virus [J]. Vet Res, 2007, 38(2): 281-97.
[3] Cavanagh D, Davis P J, Mockett A P. Amino acids
within hypervariable region 1 of avian coronavirus
IBV (Massachusetts serotype) spike glycoprotein are
associated with neutralization epitopes [J]. Virus Res,
1988, 11(2): 141-50.
[4] Ma T, Xu L, Ren M, et al. Novel genotype of
infectious bronchitis virus isolated in China [J]. Vet
Microbiol, 2019, 230: 178-86.
[5] Petzoldt D, Vogel N, Bielenberg W, et al. IB80-A
Novel Infectious Bronchitis Virus Genotype (GVIII)
[J]. Avian Dis, 2022, 66(3): 1-8.
[6] Valastro V, Holmes E C, Britton P, et al. S1 gene￾based phylogeny of infectious bronchitis virus: An
attempt to harmonize virus classification [J]. Infect
Genet Evol, 2016, 39: 349-64.
[7] Cook J K, Jackwood M, Jones R C. The long view:
40 years of infectious bronchitis research [J]. Avian
Pathol, 2012, 41(3): 239-50.
[8] Jackwood M W, Hilt D A, Lee C W, et al. Data
from 11 years of molecular typing infectious bronchitis
virus field isolates [J]. Avian Dis, 2005, 49(4): 614-8.
[9] Jackwood M W, Hilt D A, Williams S M, et
al. Molecular and serologic characterization,
pathogenicity, and protection studies with infectious
bronchitis virus field isolates from California [J].
Avian Dis, 2007, 51(2): 527-33.
[10] Jackwood M W, Jordan B J. Molecular Evolution
of Infectious Bronchitis Virus and the Emergence of
Variant Viruses Circulating in the United States [J].
Avian Dis, 2021, 65(4): 631-6.
[11] Martin E A, Brash M L, Hoyland S K, et al.
Genotyping of infectious bronchitis viruses identified
in Canada between 2000 and 2013 [J]. Avian Pathol,
2014, 43(3): 264-8.
[12] Fraga A P, Graf T, Pereira C S, et al.
Phylodynamic analysis and molecular diversity of the
avian infectious bronchitis virus of chickens in Brazil
[J]. Infect Genet Evol, 2018, 61: 77-83.
[13] Bande F, Arshad S S, Omar A R, et al. Global
distributions and strain diversity of avian infectious
bronchitis virus: a review [J]. Anim Health Res Rev,
2017, 18(1): 70-83.
[14] Cook J K. The classification of new serotypes of
infectious bronchitis virus isolated from poultry flocks
in Britain between 1981 and 1983 [J]. Avian Pathol,
1984, 13(4): 733-41.
[15] Marandino A, Perez R. Genetic and Antigenic
Diversity of Infectious Bronchitis Virus in South
America [J]. Avian Dis, 2021, 65(4): 624-30.
[16] Domanska-Blicharz K, Sajewicz-Krukowska
J, Lisowska A. New PA/1220/98-like variant of
infectious bronchitis virus in Poland [J]. Avian Pathol,
2020, 49(4): 380-8.
[17] 林绍仪 , 刘镇明 , 邝荣禄 , 等 . 鸡传染性支气管
炎肾变病型的诊断报告 [J]. 养禽与禽病防治 , 1983,
(01): 49-51.
[18] Zhao Y, Zhang H, Zhao J, et al. Evolution of
infectious bronchitis virus in China over the past two
decades [J]. J Gen Virol, 2016, 97(7): 1566-74.
[19] Chen L, Xiang B, Hong Y, et al. Phylogenetic
analysis of infectious bronchitis virus circulating in
southern China in 2016-2017 and evaluation of an
attenuated strain as a vaccine candidate [J]. Arch
Virol, 2021, 166(1): 73-81.
[20] Sun L, Tang X, Qi J, et al. Two newly isolated GVI
lineage infectious bronchitis viruses in China show
unique molecular and pathogenicity characteristics [J].
Infect Genet Evol, 2021, 94: 105006.
[21] Lian J, Wang Z, Xu Z, et al. Distribution and
molecular characterization of avian infectious
14 2026 年第 2 期
bronchitis virus in southern China [J]. Poult Sci, 2021,
100(7): 101169.
[22] Fan W, Chen J, Zhang Y, et al. Phylogenetic and
Spatiotemporal Analyses of the Complete Genome
Sequences of Avian Coronavirus Infectious Bronchitis
Virus in China During 1985-2020: Revealing Coexistence
of Multiple Transmission Chains and the Origin of LX4-
Type Virus [J]. Front Microbiol, 2022, 13: 693196.
[23] Yuan W, Lv T, Jiang W, et al. Antigenic
Characterization of Infectious Bronchitis Virus in the
South China during 2021-2022 [J]. Viruses, 2023, 15(6).
[24] Chen L, Jiang W, Wu W, et al. Insights into the
Epidemiology, Phylodynamics, and Evolutionary
Changes of Lineage GI-7 Infectious Bronchitis Virus
[J]. Transbound Emerg Dis, 2023, 2023: 9520616.
[25] Jiang L, Zhao W, Han Z, et al. Genome
characterization, antigenicity and pathogenicity of a
novel infectious bronchitis virus type isolated from
south China [J]. Infect Genet Evol, 2017, 54: 437-46.
[26] Xiao H, Yan X, Yu S, et al. A strain of avian
coronavirus of the GVII lineage capable of infecting
African green monkey kidney cell lines (Vero) and
chicken embryo fibroblast cell lines (DF-1) [J]. Poult
Sci, 2025, 104(12): 105971.
[27] Feng K, Wang F, Xue Y, et al. Epidemiology and
characterization of avian infectious bronchitis virus
strains circulating in southern China during the period
from 2013-2015 [J]. Sci Rep, 2017, 7(1): 6576.
[28] Feng K, Xue Y, Wang F, et al. Analysis of S1
gene of avian infectious bronchitis virus isolated in
southern China during 2011-2012 [J]. Virus Genes,
2014, 49(2): 292-303.
[29] Ji J, Gao Y, Chen Q, et al. Epidemiological
investigation of avian infectious bronchitis and locally
determined genotype diversity in central China: a
2016-2018 study [J]. Poult Sci, 2020, 99(6): 3001-8.
[30] Wang Y, Li Y, Yu X, et al. Epidemiological
surveillance of avian coronavirus in China in 2024 [J].
Poult Sci, 2025, 104(7): 105251.
[31] 张春阳 , 兰邹然 , 李宏梅 , 等 . 山东部分地区禽
传染性支气管炎流行病学调查 [J]. 中国家禽 , 2024,
46(06): 86-92.
[32] 刘庆伦 , 刘高升 . 聊城市阳谷县鸡传染性支气管
炎病毒感染情况调查及 S1 基因遗传进化分析 [J]. 畜
牧与饲料科学 , 2025, 46(01): 104-8.
[33] 李彬 , 韩晓青 , 刘红祥 , 等 . 2019—2021 年我国
29 个省级行政区鸡传染性支气管炎流行病学调查
[J]. 中国动物检疫 , 2022, 39(04): 19-25.
[34] 张明伟 , 赵君 , 孙璐 , 等 . 我国 2021-2023 年鸡
传染性支气管炎病毒流行趋势研究 [J]. 病毒学报 ,
2024, 40(05): 1099-104.
[35] 张小荣 , 吴艳涛 . 禽传染性支气管炎流行动态与
防控对策 [J]. 中国家禽 , 2016, 38(16): 1-5.
[36] 陈良珂 , 封柯宇 , 李鸿鑫 , 等 . 2013 ~ 2017 年鸡
传染性支气管炎病毒分子流行病学分析 [J]. 中国家
禽 , 2018, 40(15): 16-20.
[37] 赵佳 . 2018 ~ 2021 年我国部分地区鸡传染性支
气管炎病毒分子流行病学调查及其抗体检测 ELISA
方法的建立与应用 [D]. 扬州 : 扬州大学 , 2022.
[38] 何云凤 , 赵鸿洁 , 郭霄慧 , 等 . 河北地区鸡传染
性支气管炎病毒的遗传变异分析 [J]. 中国预防兽医
学报 , 2022, 44(06): 661-6.
[39] Xie Y, Fan W, Lin Z, et al. Evolution and
transmission dynamics of infectious bronchitis virus
in southern China during 2018-2023: Emergence of a
novel GI-19 subtype and re-emergence of GI-28 [J].
Poult Sci, 2025, 104(10): 105564.
[40] Xiong T, Xie H, Li L, et al. Prevalence, Genotype
Diversity, and Distinct Pathogenicity of 205
Gammacoronavirus Infectious Bronchitis Virus Isolates
in China during 2019-2023 [J]. Viruses, 2024, 16(6).
[41] Meng X, Zhang J, Wan Z, et al. Molecular
epidemiology of infectious bronchitis virus in eastern
and southern China during 2021-2023 [J]. Poult Sci,
2024, 103(8): 103939.
[42] Kong L, You R, Zhang D, et al. Infectious Bronchitis
Virus Infection Increases Pathogenicity of H9N2 Avian
Influenza Virus by Inducing Severe Inflammatory
Response [J]. Front Vet Sci, 2021, 8: 824179.
[43] 陈文平 , 康超 , 易辰阳 , 等 . 滑液囊支原体和传
染性支气管炎共感染模型的建立及初步研究 [J]. 中
国动物传染病学报 , 2024, 32(06): 8-16.
[44] Shen Y X, Li W W, Xia J, et al. Research Note:
Effects of Escherichia coli co-infection on the protective
efficacy assessment of two common infectious bronchitis
vaccines [J]. Poult Sci, 2021, 100(9): 101324.